Duckling Huddle by Charles Davis, of wood ducks in a late spring snowstorm in Smiggin Holes, New South Wales, Australia, was shortlisted. Charles Davis / PA
Ice Bed by Nima Sarikhani, of a polar bear in Norway's Svalbard archipelago, has been voted as winner of the Wildlife Photographer of the Year People's Choice Award. Nima Sarikhani / PA
The Happy Turtle by Tzahi Finkelstein, of a Balkan pond turtle with a dragonfly in Israel's Jezreel Valley, is one of the highly commended images. Tzahi Finkelstein / PA
Aurora Jellies by Audun Rikardsen, from Norway, of moon jellyfish in a fjord outside Tromso illuminated by the aurora borealis, was also highly commended. Audun Rikardsen / PA
Shared Parenting by Mark Boyd, of lions in Maasai Mara, Kenya, was highly commended. Mark Boyd / PA
Starling Murmuration by Daniel Dencescu, taken in Rome, Italy, is another highly commended image. Daniel Dencescu / PA
Swallow Over Meadow by Hermann Hirsch and Jan Lessman was shortlisted. Hermann Hirsch / Jan Lessman / PA
Bull In A Garbage Dump by Brent Stirton of a scavenging male elephant kicking over rubbish in Tissamaharama, Sri Lanka, was shortlisted. Brent Stirton / PA
Curiosity by Gerald Hinde, of lions in South Africa's Greater Kruger National Park, was shortlisted. Gerald Hinde / PA
Homecoming by Dvir Barkay, of a pygmy round-eared bat returning to its home in a termite nest in Costa Rica, was shortlisted. Dvir Barkay / PA
Opportunity Fox by Matt Maran, of a young red fox in London, was shortlisted. Matt Maran / PA
Looking At Me, Looking At You by John E Marriott, of a grizzly bear in the Chilko River, British Columbia, Canada, was shortlisted. John E. Marriott / PA
Troublemaker by Stefan Christmann, of an Adelie penguin approaching an emperor penguin and its chick during feeding time in Antarctica's Atka Bay, was shortlisted. Stefan Christmann / PA
Snowshoes by Deena Sveinsson, of a snowshoe hare in the Rocky Mountain National Park, US,was shortlisted. Deena Sveinsson / PA
Neighbourhood Dispute by Ofer Levy, of a mudskipper fiercely defending its territory from a trespassing crab in Roebuck Bay, Australia, was shortlisted. Ofer Levy / PA
The Grassland Geladas by Marco Gaiotti, of Gelada monkeys in the Simien Mountains of Ethiopia, was shortlisted. Marco Gaiotti / PA
Missed Sip of Milk by Karim Iliya, of a humpback whale calf missing some of its mother's milk off the coast of Rurutu, French Polynesia, was shortlisted. Karim Iliya / PA
Hope by Roberto Garcia-Roa, of a rescued chimpanzee looking out of its enclosure at the Chimpanzee Conservation Centre in the Republic of Guinea, was shortlisted. Roberto Garcia-Roa / PA
Incoming Cuckoo Wasp by Frank Deschandol, of a cuckoo wasp trying to enter a mason bee's clay burrow near Montpellier, France, was shortlisted. Frank Deschandol / PA
A Rare Sight by Axel Gomille, of an Ethiopian wolf in Ethiopia's Bale Mountains National Park, was shortlisted. Axel Gomille / PA
Tender Touch by Andy Parkinson, of two courting mountain hares in the Monadhliath Mountains in Scotland, was shortlisted. Andy Parkinson / PA
Rubbish Drinks by Claire Waring, of a Celebes crested macaque investigating a plastic bottle at the edge of Tangkoko Batuangus Nature Reserve, Indonesia, was shortlisted. Claire Waring / PA
Fashion Victims by Britta Jaschinski, of confiscated coats made from endangered big cat skins in Hamburg, was shortlisted. Britta Jaschinski / PA
Autumn Glow by Uge Fuertes Sanz, of bullrushes and quaking aspens by the Cabriel River in the Sierra de Albarracin, Spain, was shortlisted. Uge Fuertes Sanz / PA
Duckling Huddle by Charles Davis, of wood ducks in a late spring snowstorm in Smiggin Holes, New South Wales, Australia, was shortlisted. Charles Davis / PA
Ice Bed by Nima Sarikhani, of a polar bear in Norway's Svalbard archipelago, has been voted as winner of the Wildlife Photographer of the Year People's Choice Award. Nima Sarikhani / PA
The Happy Turtle by Tzahi Finkelstein, of a Balkan pond turtle with a dragonfly in Israel's Jezreel Valley, is one of the highly commended images. Tzahi Finkelstein / PA
Aurora Jellies by Audun Rikardsen, from Norway, of moon jellyfish in a fjord outside Tromso illuminated by the aurora borealis, was also highly commended. Audun Rikardsen / PA
Shared Parenting by Mark Boyd, of lions in Maasai Mara, Kenya, was highly commended. Mark Boyd / PA
Starling Murmuration by Daniel Dencescu, taken in Rome, Italy, is another highly commended image. Daniel Dencescu / PA
Swallow Over Meadow by Hermann Hirsch and Jan Lessman was shortlisted. Hermann Hirsch / Jan Lessman / PA
Bull In A Garbage Dump by Brent Stirton of a scavenging male elephant kicking over rubbish in Tissamaharama, Sri Lanka, was shortlisted. Brent Stirton / PA
Curiosity by Gerald Hinde, of lions in South Africa's Greater Kruger National Park, was shortlisted. Gerald Hinde / PA
Homecoming by Dvir Barkay, of a pygmy round-eared bat returning to its home in a termite nest in Costa Rica, was shortlisted. Dvir Barkay / PA
Opportunity Fox by Matt Maran, of a young red fox in London, was shortlisted. Matt Maran / PA
Looking At Me, Looking At You by John E Marriott, of a grizzly bear in the Chilko River, British Columbia, Canada, was shortlisted. John E. Marriott / PA
Troublemaker by Stefan Christmann, of an Adelie penguin approaching an emperor penguin and its chick during feeding time in Antarctica's Atka Bay, was shortlisted. Stefan Christmann / PA
Snowshoes by Deena Sveinsson, of a snowshoe hare in the Rocky Mountain National Park, US,was shortlisted. Deena Sveinsson / PA
Neighbourhood Dispute by Ofer Levy, of a mudskipper fiercely defending its territory from a trespassing crab in Roebuck Bay, Australia, was shortlisted. Ofer Levy / PA
The Grassland Geladas by Marco Gaiotti, of Gelada monkeys in the Simien Mountains of Ethiopia, was shortlisted. Marco Gaiotti / PA
Missed Sip of Milk by Karim Iliya, of a humpback whale calf missing some of its mother's milk off the coast of Rurutu, French Polynesia, was shortlisted. Karim Iliya / PA
Hope by Roberto Garcia-Roa, of a rescued chimpanzee looking out of its enclosure at the Chimpanzee Conservation Centre in the Republic of Guinea, was shortlisted. Roberto Garcia-Roa / PA
Incoming Cuckoo Wasp by Frank Deschandol, of a cuckoo wasp trying to enter a mason bee's clay burrow near Montpellier, France, was shortlisted. Frank Deschandol / PA
A Rare Sight by Axel Gomille, of an Ethiopian wolf in Ethiopia's Bale Mountains National Park, was shortlisted. Axel Gomille / PA
Tender Touch by Andy Parkinson, of two courting mountain hares in the Monadhliath Mountains in Scotland, was shortlisted. Andy Parkinson / PA
Rubbish Drinks by Claire Waring, of a Celebes crested macaque investigating a plastic bottle at the edge of Tangkoko Batuangus Nature Reserve, Indonesia, was shortlisted. Claire Waring / PA
Fashion Victims by Britta Jaschinski, of confiscated coats made from endangered big cat skins in Hamburg, was shortlisted. Britta Jaschinski / PA
Autumn Glow by Uge Fuertes Sanz, of bullrushes and quaking aspens by the Cabriel River in the Sierra de Albarracin, Spain, was shortlisted. Uge Fuertes Sanz / PA
Duckling Huddle by Charles Davis, of wood ducks in a late spring snowstorm in Smiggin Holes, New South Wales, Australia, was shortlisted. Charles Davis / PA